To all the people talking about the lack of transitional fossils, whats the deal? We have plenty. There's even a section there Which I'll quote about elephants specifically (since it came up ) (but I'd recommend just reading the link, formatting doesn't work well on this forum)
Originally posted by http://www.talkorigins.org/faqs/faq-...t2b.html#eleph
Elephants
* Minchenella or a similar condylarth (late Paleocene) -- Known only from lower jaws. Has a distinctive broadened shelf on the third molar. The most plausible ancestor of the embrithopods & anthracobunids.
* Phenacolophus (late Paleocene or early Eocene) -- An early embrithopod (very early, slightly elephant-like condylarths), thought to be the stem-group of all elephants.
* Pilgrimella (early Eocene) -- An anthracobunid (early proto-elephant condylarth), with massive molar cusps aligned in two transverse ridges.
* Unnamed species of proto-elephant (early Eocene) -- Discovered recently in Algeria. Had slightly enlarged upper incisors (the beginnings of tusks), and various tooth reductions. Still had \"normal\" molars instead of the strange multi-layered molars of modern elephants. Had the high forehead and pneumatized skull bones of later elephants, and was clearly a heavy-boned, slow animal. Only one meter tall.
* Moeritherium, Numidotherium, Barytherium (early-mid Eocene) -- A group of three similar very early elephants. It is unclear which of the three came first. Pig-sized with stout legs, broad spreading feet and flat hooves. Elephantish face with the eye set far forward & a very deep jaw. Second incisors enlarged into short tusks, in upper and lower jaws; little first incisors still present; loss of some teeth. No trunk.
* Paleomastodon, Phiomia (early Oligocene) -- The first \"mastodonts\", a medium-sized animals with a trunk, long lower jaws, and short upper and lower tusks. Lost first incisors and canines. Molars still have heavy rounded cusps, with enamel bands becoming irregular. Phiomia was up to eight feet tall.
GAP: Here's that Oligocene gap again. No elephant fossils at all for several million years.
* Gomphotherium (early Miocene) -- Basically a large edition of Phiomia, with tooth enamel bands becoming very irregular. Two long rows cusps on teeth became cross- crests when worn down. Gave rise to several families of elephant- relatives that spread all over the world. From here on the elephant lineages are known to the species level.
* The mastodon lineage split off here, becoming more adapted to a forest browser niche, and going through Miomastodon (Miocene) and Pliomastodon (Pliocene), to Mastodon (or \"Mammut\", Pleistocene).
Meanwhile, the elephant lineage became still larger, adapting to a savannah/steppe grazer niche:
* Stegotetrabelodon (late Miocene) -- One of the first of the \"true\" elephants, but still had two long rows of cross-crests, functional premolars, and lower tusks. Other early Miocene genera show compression of the molar cusps into plates (a modern feature ), with exactly as many plates as there were cusps. Molars start erupting from front to back, actually moving forward in the jaw throughout life.
* Primelephas (latest Miocene) -- Short lower jaw makes it look like an elephant now. Reduction & loss of premolars. Very numerous plates on the molars, now; we're now at the modern elephants' bizarre system of one enormous multi-layered molar being functional at a time, moving forward in the jaw.
* Primelephas gomphotheroides (mid-Pliocene) -- A later species that split into three lineages, Loxodonta, Elephas, and Mammuthus:
1. Loxodonta adaurora (5 Ma). Gave rise to the modern African elephant Loxodonta africana about 3.5 Ma.
2. Elephas ekorensis (5 Ma), an early Asian elephant with rather primitive molars, clearly derived directly from P. gomphotheroides. Led directly to:
o Elephas recki, which sent off one side branch, E. hydrusicus, at 3.8 Ma, and then continued changing on its own until it became E. iolensis.
o Elephas maximus, the modern Asian elephant, clearly derived from
o E. hysudricus. Strikingly similar to young E. hysudricus animals. Possibly a case of neoteny (in which \"new\" traits are simply juvenile features retained into adulthood).
3. Mammuthus meridionalis, clearly derived from P. gomphotheroides. Spread around the northern hemisphere. In Europe, led to M. armeniacus/trogontherii, and then to M. primigenius. In North America, led to M. imperator and then M. columbi.
The Pleistocene record for elephants is very good. In general, after the earliest forms of the three modern genera appeared, they show very smooth, continuous evolution with almost half of the speciation events preserved in fossils. For instance, Carroll (1988) says: \"Within the genus Elephas, species demonstrate continuous change over a period of 4.5 million years. ...the elephants provide excellent evidence of significant morphological change within species, through species within genera, and through genera within a family....\"
Species-species transitions among the elephants:
* Maglio (1973) studied Pleistocene elephants closely. Overall, Maglio showed that at least 7 of the 17 Quaternary elephant species arose through smooth anagenesis transitions from their ancestors. For example, he said that Elephas recki \"can be traced through a progressive series of stages...These stages pass almost imperceptibly into each other....In the late Pleistocene a more progressive elephant appears which I retain as a distinct species, E. iolensis, only as a matter of convenience. Although as a group, material referred to E. iolensis is distinct from that of E. recki, some intermediate specimens are known, and E. iolensis seems to represent a very progressive, terminal stage in the E. recki specific lineage.\"
* Maglio also documented very smooth transitions between three Eurasian mammoth species: Mammuthus meridionalis --> M. armeniacus (or M. trogontherii) --> M. primigenius.
* Lister (1993) reanalyzed mammoth teeth and confirmed Maglio's scheme of gradual evolution in European mammoths, and found evidence for gradual transitions in the North American mammoths too.
Not being an evolutionary biologist I can't really present my own case for these transitional fossils but if you're interested enough to follow the thread this far, you'll probably like to follow it a bit further to that link as well . It also has a nice explenation for why gaps occur.
-spoon
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